A quaint midrash has been applied to this verse, viewing the lulav (palm frond) as symbolic of the human spine, the hadas (myrtle branch) symbolic of the eye, the arava (willow branch) symbolic of the lips, and the etrog (citron) symbolic of the heart.

When all four species are taken together in the festival "bouquet," they represent all the essential parts of the human body uniting to praise God.

“A land of wheat and barley, of vines, figs, and pomegranates, a land of olive oil and of (date) honey…”
Deuteronomy 8:8